The genus Brachiaria includes the most widely used
forage grasses in the American tropics. Traditional commercial
cultivars are direct selections from ex situ germplasm collections
of species of African origin. Except for the diploid sexual
B. ruziziensis, existing Brachiaria cultivars
(B. brizantha, B. decumbens, and B. humidicola)
are polyploid apomicts. Genetic improvement projects were
contemplated at least from the early 1970s to rectify the
deficiencies of the existing commercial cultivars. However,
apomictic reproduction (asexual reproduction through seeds)
and ploidy differences among and within species blocked effective
plant breeding until the mid-1980s, when a tetraploid sexual
biotype of B. ruziziensis was developed in Belgium.
In early 1988, this invaluable germplasm reached CIAT through
our Brazilian colleague Dr. Cacilda do Valle, geneticist and
plant breeder at EMBRAPA's National Center for Research on
Beef Cattle in Campo Grande, Mato Grosso do Sul.
The cross-compatible, sexual tetraploid allowed recombination
of genes that previously were reproductively inaccessible
in tetraploid apomicts. The first experimental hybrids were
tested in a small field trial in Colombia in 1989. From this
modest beginning, a full-scale genetic improvement program
developed over the succeeding years.
A Mexican-based forage seed company, Semillas Papalotla,
multiplied and commercialized the first hybrid apomictic Brachiaria
cultivar under the name Mulato. Its prime attributes are high
yield and feed quality. However, Mulato's commercial success
was limited by its low seed yield potential. A second hybrid,
called Mulato II was released to the commercial market in
2005. Mulato II has better spittlebug resistance than Mulato
and is more drought tolerant. The limited seed availability
of Mulato II in 2005 has been overcome and seed is commercially
available in most countries.
Continuing cycles of hybridization and selection seek to
produce hybrids with enhanced expression of spittlebug resistance,
forage yield and quality, seed yield, and other desirable
attributes.
Related publications:
Miles,
J.W.; do Valle, C.B.; Rao, I.M.; Euclides, V.P.B. 2004. Brachiaria
grasses. In: Sollenberger, L.E.; Moser, L.; Burson, B. (eds.).
Warm-season (C4) grasses. Agron. Monogr. 45. ASA, CSSA, SSSA,
Madison, WI, USA. p. 745-783.
Miles,
J.W.; Cardona, C.; Sotelo, G. 2006. Recurrent selection in
a synthetic brachiariagrass population improves resistance
to three spittlebug species. Crop Sci. 46:1088-1093.
Responsible: John
W. Miles
Collaborators: Semillas Papalotla
(e-mail: mexico@grupopapalotla.com;
Web: www.grupopapalotla.com)

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